The spines arise in two series from the areoles. The radial spines, 9—11 in the pictures below, are shorter and narrower than the central spines and tend to spread parallel with the surface outline of the tubercule-covered stem. The larger, wide and flattened central spines are fewer in number, 4 below, and project outward from the aereole; the lowest of these (most distant from the stem apex) with a hook.
The two photo series below show the original photo detail containing the spine set of a single aereole, on the left; and a colored(ized) schematization on the right: radial spines in yellow, the hook spine in purple, other central spines in red.
The surface of the spines is covered with tiny hairs, as may
be seen below when backlit.