To Berberis Home

A 5-foliolate "B. trifoliolata"

by Bob Harms  email-here

Trifoliate bush with 5 foliolate leaves (Jan. 2005)

One bush with nearly all trifoliate leaves has repeatedly developed an extra leaf pair, at times virtually merged with the node of the topmost leaflets.


Uniquely merged leaflet nodes

This formation, unique among hundreds of bushes surveyed, would appear to reflect introgressive hybridization resulting in an almost total B. trifoliolata gene pool, additionally supporated by its 4-5-spinose leaflet margins. But this bush otherwise lacked phenological and morphological indicators of hybridization, such as dentate filaments.


5 foliolate leaf with rachis separating the nodes for the lower leaflet pair (October 2005; one leaflet has fallen off)